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Acquisition - Schedule of Estimated Fair Values of Acquired (Details) link:presentationLink link:calculationLink link:definitionLink 9955527 - Disclosure - License, Collaboration and Success Payment Agreements - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - License, Collaboration and Success Payment Agreements - Schedule of Aggregate Potential Success Payments (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Impairment of Long-Lived Assets (Details) link:presentationLink link:calculationLink link:definitionLink 9955530 - Disclosure - Cash Equivalents and Marketable Securities - Schedule of Fair Value and Amortized Cost of Cash Equivalents and Marketable Securities (Details) link:presentationLink link:calculationLink link:definitionLink 9955531 - Disclosure - Cash Equivalents and Marketable Securities - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Other Investments (Details) link:presentationLink link:calculationLink link:definitionLink 9955533 - Disclosure - Fair Value Measurements - Schedule of Financial Assets and Liabilities Measured at Fair Value on Recurring Basis (Details) link:presentationLink link:calculationLink link:definitionLink 9955534 - Disclosure - Fair Value Measurements - Schedule of Estimated Fair Value of Success Payment Liability Assumptions (Details) link:presentationLink link:calculationLink link:definitionLink 9955535 - Disclosure - Fair Value Measurements - Schedule of Changes in Estimated Fair Value of Financial Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Fair Value Measurements - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955537 - Disclosure - Property and Equipment, Net - Schedule of Property and Equipment, Net (Details) link:presentationLink link:calculationLink link:definitionLink 9955538 - 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